Rafaela Sayuri Takeshita

PhD Candidate (D3)

Social Systems Evolution


I am interested in behavioral and physiological responses to social and environmental changes in nonhuman primates. My research involves non-invasive hormonal analysis and behavioral observation to:

  • Investigate the impact of environment on primate welfare and reproductive fitness
  • Monitor reproductive state and stress-related responses in captive and wild populations
  • Associate hormonal secretion pattern with the evolution of social systems in different primate species


2013 – current     D.Sc. Primatology, Graduate School of Science, Kyoto University.

2011 – 2013        M.Sc. Primatology, Graduate School of Science, Kyoto University. Thesis: “Non-invasive analysis of adrenal hormones in female japanese macaques (Macaca fuscata)”

2006 – 2011        B.Sc. Veterinary Medicine, Instituto da Saúde e Produção Animal, Universidade Federal Rural da Amazônia. Thesis: “Avaliação hematológica, hepatica e renal em Aotus azarai infulatus” (Hematological, hepatic, and renal evaluation in Aotus azarai infulatus).

Selected Publications

Takeshita RSC, Huffman MA, Kinoshita K, Bercovitch FB (2017). Effect of castration on social behavior and hormones in male Japanese macaques (Macaca fuscata). Physiology & Behavior 181:43-50 (doi: 10.1016/j.physbeh.2017.09.006).

Takeshita RSC, Huffman MA, Mouri K, Shimizu K, Bercovitch FB (2016). Dead or alive? Predicting fetal loss in Japanese macaques (Macaca fuscata) by fecal metabolites. Animal Reproduction Science 175: 33-38 (doi: 10.1016/j.anireprosci.2016.10.006).

Takeshita RSC, Huffman MA, Bercovitch FB (2015). Non-invasive analysis of adrenal hormones in Japanese macaques. Saarbrücken, Germany: LAP Lambert Academic. 83p.

Takeshita RSC, Bercovitch FB, Huffman MA, Mouri K, Garcia C, Rigaill L, Shimizu K (2014). Environmental, biological, and social factors influencing fecal adrenal steroid concentrations in female Japanese macaques (Macaca fuscata). American Journal of Primatology 76 (11):1084-1093.

Takeshita RSC, Huffman MA, Bercovitch FB, Mouri K, Shimizu K (2013). The influence of age and season on fecal dehydroepiandrosterone-sulfate (DHEAS) concentrations in Japanese macaques (Macaca fuscata). General and Comparative Endocrinology 191: 39-43.

Takeshita RSC, Monteiro FOB, Lins FLML, Silva GA, Faturi C, Coutinho LN, Monteiro MVB, Kugelmeier T, Castro PHG, Muniz JAPC (2011). Hematological, hepatic, and renal evaluation in Aotus azarai infulatus. Journal of Medical Primatology 40: 104 – 110.


Social systems evolution section

Primate Research Institute, Kyoto University
41-2 Kanrin, Inuyama, Aichi
Japan 484-8506